Four million years after the Autobots' spacecraft - The Ark, crashlanded on the once lifeless planet of Earth, The Autobot and Decepticon passengers have been revived by Auntie, the Ark's main computer. Each Transformer has been given the ability to alter their form into Earth vehicles, weapons, and military machinery. The Decepticon forces, commanded by the malevolent Megatron, decide to flee the crippled spacecraft to recover and recharge before they continue with their mission, to destroy the Autobots once and for all!